A few fish released into an empty lake multiply quickly at first, because food and space are plentiful. As the lake fills, each new fish has less to eat, and growth slows until the population levels off. An exponential model gets the beginning right and the end badly wrong. The logistic model describes both stages with three parameters, each with a clear meaning.
Key terms
Logistic function
Definition A logistic function is a function of the form
with constants , and . Its domain is all real numbers and its range is the open interval . It is increasing, and its graph has the horizontal asymptote as and as . The graph has the shape of a stretched letter S. Books use different letters for these constants, and the calculus lesson on logistic growth writes in place of and in place of .
In plain words A logistic function starts near zero, climbs, and then flattens out under a ceiling it never touches.
Example A lake is stocked with fish of a new species, and the fish population years later is modeled by
Rounded to whole fish, the model gives these values.
| in years | ||||||
|---|---|---|---|---|---|---|
Positive parameters
Definition In all three parameters must be positive. The starting value is
With , the condition holds exactly when , and makes increasing. Under these conditions the denominator is greater than for every , so is defined everywhere and stays between and . If instead while , the denominator is zero at and the graph has a vertical asymptote there. If but , the curve falls toward as grows.
In plain words Positive parameters guarantee the S shape. The ceiling is positive, the start is between zero and the ceiling, and the curve goes up.
Example For the lake, and , so
If a survey had instead counted fish at the start, more than the lake can support, the formula would give
With that negative and , the curve falls from toward for instead of rising in an S. A start above the capacity is outside the definition above, which describes growth from below.
Carrying capacity
Definition In a logistic model the parameter is the carrying capacity, the size the population approaches over time. Because , every value satisfies , and because as ,
So the line is a horizontal asymptote of the graph. It is approached but never reached.
In plain words The carrying capacity is the most the environment can hold in the long run. The model gets as close to it as you like but never gets there.
Example For the lake the carrying capacity is fish. After years the model gives about fish, and after years
still below .
Early growth
Definition Multiplying by and rearranging gives, for every , the exact identity
While is small compared with , the factor in parentheses is close to , so
Early logistic growth is therefore nearly exponential with continuous growth rate . The exponential formula is too large by the factor , which grows as approaches .
In plain words At the start the ceiling is far away and barely matters, so the population grows like an exponential function. The approximation only fails once the population is a noticeable fraction of the capacity.
Example For the lake, .
The two columns stay within about percent for four years. By year the exponential guess is more than percent too large.
Saturation
Definition Saturation is the late stage of logistic growth, when is close to and growth has nearly stopped. The remaining gap is
which is positive for every and approaches as . Once is small the gap is approximately , so it shrinks by about the factor with each unit of time.
In plain words Near the ceiling the population creeps upward, and each year closes about the same fraction of the remaining gap.
Example Here is the gap for the lake every two years. Over two years the factor is .
Each entry is close to times the one before it.
Symmetry about the midpoint
Definition The midpoint of a logistic curve is the point with
Since , the function can be written as
and . The graph is symmetric about the midpoint, which means
To see this, put . Then
which add to . The curve is also steepest at the midpoint, where it changes from bending upward to bending downward. Calculus proves that fact, and here it is taken as known.
In plain words Turn the graph half a turn about its midpoint and it lands on itself. The growth that happens just before the midpoint is mirrored by the growth just after it.
Example For the lake, years, when the population is . Four years on either side,
which add to . The population gains about fish in the four years before the midpoint and the same number in the four years after.
Model interpretation without differential equations
Definition In calculus the logistic function comes from an equation about its rate of change, a differential equation. Without that equation, a logistic model can still be read entirely from its formula. The carrying capacity is , the starting value is , the early continuous growth rate is , and is the time of half capacity and fastest growth. For any level with , solving with logarithms gives the unique time at which the model reaches ,
The levels and are never reached. The model also rests on assumptions that real populations can break. The capacity stays fixed, growth is smooth with no seasons, harvests or disasters, and the rise is exactly symmetric about the midpoint.
In plain words The formula and logarithms answer every question the model can answer. Questions outside its assumptions, like what happens if the lake shrinks in a drought, it cannot answer at all.
Example The lake model reaches fish when
Asking when it reaches fish has no answer, since the model only approaches that level. A real lake is also unlikely to settle at exactly . Its population usually rises and falls around the capacity from year to year, which this smooth curve cannot show.
Check yourself
- For the model , find the carrying capacity, the starting value and the time of the midpoint.
Show the answer
The carrying capacity is . The starting value is . The midpoint time is
when the value is .
- For the same model, when does reach ?
Show the answer
Setting gives
So and . Taking natural logarithms gives , so
- A logistic model has carrying capacity and midpoint time . Three time units before the midpoint its value is . What is its value three time units after the midpoint?
Show the answer
By symmetry about the midpoint, , so .